In the week Owen Farrell and Billy Vunipola were issued with bans that rule them out of the opener against Argentina, head coach Steve Borthwick has now lost a first-choice wing.
Watson sustained the damage in Saturday's 29-10 rout by Ireland and has been replaced in the 33-man World Cup squad by Jonny May, who starts tomorrow's clash with Fiji.
It is a savage blow for the Lions three-quarter who had previously battled back from long-term knee and Achilles injuries and was poised to start the Pumas showdown on September 9.
"In the game against Ireland, Anthony Watson suffered an injury to his calf which means he can't play for a number of weeks," Borthwick said.
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