A legendary figure for the Lions, Harris scored a club-record 138 goals across two spells as a player before embarking on a successful four-year stint as manager that yielded promotion to the Championship in 2017.
Exhausted, he stepped down two years later and has since spent time in charge at Cardiff, Gillingham and, most recently, Cambridge.
Harris, though, has always maintained that he'd be open to a return and when the disastrous 19game reign of rookie boss Joe Edwards was terminated on Wednesday, the 46-year-old quit the U's to answer Millwall's SOS.
"I was always very clear that I'd be here for the club when needed," said Harris, who started the season at Gillingham and joined Cambridge in December, thus achieving the unusual feat of managing at all three levels of the EFL within a single campaign.
Ready
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LUKE BOOSTED BY WALES ACE LIAM
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MATT'S DELIGHT OVER RECORD
WYCOMBE boss Wanderers Matt Bloomfield hailed his history-makers after victory at Lincoln City.
TON-UP IAN ENJOYS HIS SPECIAL WIN
AN Evatt described his 100th win as Bolton boss as very special, although he needed Aaron Collins' injurytime goal to bring up the three figures.
DONS LACK A CLINICAL EDGE AS SADDLERS GRAB POINTS
NATHAN Lowe's strike inspired promotion-chasing Walsall to victory over AFC Wimbledon at the Cherry Red Records Stadium.
SUB GRIMES IS IN-SPIRED TO HEAD THE WINNER
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COBBLERS TEST IN TROPHY FOR POSH
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HORNETS REMINDED OF GRAY'S BRILLIANCE
WAYNE Rooney expressed his delight as Plymouth extended their unbeaten league home run to six after Andre Gray's last-gasp leveller.
MARTI DOUBTING JOB AFTER DRAW
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Rohl confident Owls
DANNY Rohl said getting results is the only thing missing as his Owls were held at home by Cardiff.