The Hammers are expected to announce former Brighton and Chelsea boss Graham Potter as the Spaniard's replacement in the next 24 hours.
The PA news agency understands talks with Potter were at an advanced stage on Tuesday night, with a few contract details still to be ironed out.
Lopetegui took training as usual on Tuesday and reported for work yesterday to be officially told his spell in charge was over.
"West Ham United can confirm that head coach Julen Lopetegui has today left the club.
"The first half of the 2024/25 season has not aligned with the club's ambitions and the club has therefore taken action in line with its objectives.
"The club can confirm that assistant head coach Pablo Sanz, head of performance Oscar Caro, head analyst Juan Vicente Peinado, fitness coach Borja De Alba and technical coach Edu Rubio have also left with immediate effect."
"The board would like to thank Julen and his staff for their hard work during their time with the Hammers and wish them every success for the future.
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